
Classification
Commonly known as Ant-loving
Beetles, Short-Winged Mold Beetles, or simply, pselaphids).
Until recently, they
were classified in their own family Pselaphidae (see Newton &
Thayer 1995). You will still see many references to them under
this name.
Diversity:
There are 712 species in the USA, and over 8,000 worldwide. To
give you an idea how big this is there are roughly 4,000 species
of mammals, 6,000 species of reptiles, and 8,000 species of ants.
Beetles are the most diverse group of organisms, with roughly
290,000 species. They are found all over the world.
Description:Pselaphids are small beetles, about 0.7mm to 4.5mm,
or from the
size of a small mosquito to the size of a grain of rice. They
all tend to be straw colored (see the pictures below). In the
wild they look a little like slow-moving ants. More technically,
like rove beetles (Staphylinidae), their elytra are short exposing
much of the upper surface of the abdomen, but they are different
from rove beetles in that their abdomens are usually not flexible,
their antennae are clubbed, and their abdomens tend to be broader
than their heads. For further help in identifying pselaphids and
other beetle families see Peterson's Field Guide to Beetles. For
a more technical key to the beetles see the standard entomological
textbook Introduction to Insects.
Pselaphids are very distinctive,
but it can be difficult to identify them to the genus level. A
very powerful hand lens, jeweler's visor, or best of all, a dissecting
microscope will be helpful in identification. The best key to
the genera of the United States was made by Donald S. Chandler
and appears in the Soil Biology Guide. There are other good keys
such as Arnett's Beetles of the United States.
If you're interested
in knowing what species it is, there are keys to many genera.
If you contact me, I will be glad to tell you if there is a key
for the genera you are interested. Better yet, you can send me
the specimens, I always love to see new specimens. If it's an
"easy one' I might be able to identify it, otherwise I would
send it on to one of two experts on pselaphids, Dr. Chris Carlton
or Dr. Don Chandler. You may wish to contact them directly.
Natural History:
The natural history of Pselaphids is poorly understood, but they
can be divided into species that live with ants and free-living
species. This is actually a continuum, with some species occasionally
being found with ants. The species that live with ants are somehow
able to live within the colony without being attacked as any other
insect would be. They may fool the ants into thinking they are
other ants, or just some inert matter or a dead insect. The ants
may feed the beetles directly, as they would any other ant in
the colony, or the pselaphids may eat the brood of the ants. I
find the interaction of pselaphids with ants very interesting,
and it is the subject of my research.
Many other pselaphids
are free living, occurring in a variety of habitats that share
a few characteristics, namely, they are very moist and they contain
decaying plant material. Many species are found in the leaf litter,
but they also occur in rotten logs, at the bases of grass clumps,
and under boards and rocks. Some species are specialized to live
in caves, on mountains, in bogs, on the beach, and in the desert.
All free-living pselaphids
eat other smaller animals, such as small insects and their larvae,
worms, and mites. Actually, we don't know exactly what most pselaphid
species eat, but judging from their similar mouthparts (designed
for catching prey, not grinding plants or fungus) and from what
we know a few of them eat, we can be pretty sure that the vast
majority eat small animals.
Abundance:
Unlike insects like mosquitoes or aphids, pselaphids are not abundant.
They are small and they live in hidden habitats, thus even people
who collect insects do not often encounter them. There are some
species that are amazingly scarce. They are known from only one
or a few specimens and nothing more. Many are attracted to lights,
as moths are, but we don't know where they live in the wild.
Life History:
Here is where our lack of knowledge of these beetles really shows.
For the over 8,000 species known in the world, only about 12 larvae
have been described. Many people have looked very hard for these
larvae and not found any. Are we looking in the wrong places?
Are the larvae very fast-developing? We don't know the answers.
To date, no one has uncovered the complete life history of any
pselaphid.
Collecting:
No matter where you live, you are probably just a short trip away
from a spot where pselaphids live, they may even live in that
vacant lot across the street. But as they are small, scarce, and
live in hidden places, they can be hard to collect. You rarely
just see them in the wild. Here are a few collecting methods.
Hand Collecting:
This is the most time consuming, least productive, yet most rewarding
method. When you find a pselaphid in the wild, you have a lot
of information about where they are found naturally, that you
might not get with other methods. Also I like to try and keep
them alive and see what they eat and how they behave. Most likely,
any observations on behavior or diet you make have never been
made before and are valuable scientific information. To find them
by hand you need a keen eye. Watch for slow moving, ant-like insects
in the following places: in rotten logs, under boards in fields,
under rocks (on their underside too) and in ant nests. Look carefully,
as some pselaphids 'play dead' when first disturbed. Looking in
leaf litter is too hard, although if you sift the litter through
¼ or 1/8 inch hardware cloth first, you might find something.
Berlese (burr-LAY-zee)
funnel: This is the most productive method of finding pselaphids.
To do this, first collect nice moist leaf litter (not sopping
wet), and sift it through a ¼ or 1/8 hardware cloth sifter.
This will give you a pile of sawdust-like material that may contain
a lot of insects and other arthropods running around. You put
this in a berlese funnel and let it 'run' for a few nights. For
a detailed description on what a berlese funnel is and how to
make one see either Peterson's Field Guide to Beetles or Introduction
to Insects. Essentially all it is is a funnel with a screen in
the middle, and the sifted litter is placed on the screen and
a light bulb is placed above. The heat from the light bulb dries
out the litter and drives the arthropods out the bottom of the
litter, down the funnel, to a dish of alcohol below.
Culture:
We know very little about what these beetles eat, how they mate
and raise their young. It is impractical to observe these creatures
in the wild, so raising them in captivity is the only option left.
Just about any observation you can make on a pselaphid species
living near you is likely to be new to science. They can be rather
easy to keep in captivity.
Housing: Pselaphids,
more than anything else, require high humidity. They can be easily
kept in a small container, such as a baby-food jar, plastic cottage
cheese container, or small plastic petri dishes if you can get
them.

